Published: Ann. Hist.-Nat. Mus. Natl. Hung. 70: 103 (1978)
Taxonomic status: Heterotypic synonym of H. velatum
Is basionym of: Hebeloma collariatum f. psammicola (Bohus) Bohus (1995); Hebeloma subcaespitosum var. psammicola (Bohus) Bohus [as “psammocolum”] (1983) ["1982"]
Types: HUNGARY: Between Orkeny and Tatarszentgyorgy (approx. 47.1°N, 19.38°E, alt. approx. 120 m a.s.l.) on bare, sandy soil under Populus sp., 11 Nov. 1976, M. Babos, A. Friesz, det: G. Bohus (Holotype. herbarium acc. no. BP 96.935, HJB1000053).
Original diagnosis: Pileus 3-6.5 cm, carnosulus, convexus, deinde expansus, saepe umbonatus, brunneolo-ochraceus, ochraceo-brunneus, testaceo-pallens, margine pallidior, primo velo obtectus, deinde circa marginem maculis e fibrillis albidis, demum saepe glabratus; viscosus. - Lamellulae 4-8 mm latae, demum ventricosae, sicci, e pallidis brunneolo-ochraceae, argillaceo-brunneae, adnatae-emarginatae, acie pallido. Stipes ± subcavus, 5-8 x 0.7-1.2 cm, aequalis, basi fusiformi-subradicatus, curvus, apice albidus, deorsum pallide ochraceus, basi ochraceo-brunneus, fibrillis cortinae obtectus, ad dimidium in arenam demersus, basi cum bulbo ex arena non raro magno. Caro albida in pileo at parte in stipite, dearsum in stipite ± brunnea, odore debili. - Sporae 10-12.5 x 6-6.8 μm, ellipsoideae, obtusae, subleves vel subtliter verrucosae, pallide ochraceae. Cheilocystidiae cylindraceae, 40-70 x 5- 8 μm, numerosissimae. - In sabulo vasto.
English translation: Pileus 3–6.5 cm, fleshy, convex then expanding, often umbonate, brownochre, ochre-brown, or pale brick-coloured with paler margin, first covered with veil, then with white patches and fibrils of veil, later often glabrous, viscid. Lamellae 4–8 mm broad, then ventricose, dry, first pale brown-ochre then clay-coloured brown, adnate-emarginate, with paler edge. Stipe slightly hollow, 5–8 × 0.7–1.2 cm, equal, with fusiform-rooting base, curved, with white apex, downwards pale ochre, ochre-brown at base, covered with fibrillose cortina remnants, up to halfway buried in sand, not rarely with large basal bulb often enlarged with sand. Context white in pileus and part of stipe, brownish in lower part of stipe. Smell weak. Spores 10–12.5 × 6–6.8 μm, ellipsoid, blunt, almost smooth or subtly verrucose, pale ochraceous. Cheilocystidia cylindrical, 40–70 × 5–8 μm, very numerous. In bare sand.
Commentary: We have studied the holotype morphologically and also obtained an ITS sequence from the material. This is: = Hebeloma dunense L. Corb. & R. Heim.
All current species: aanenii | adherens | aestivale | albidulum | alboerumpens | albomarginatum | album | alpinicola | alpinum | ambustiterranum | aminophilum | ammophilum | angelesiense | angustilamellatum | anthracophilum | arcticum | asperosporum | aurantioumbrinum | australe | avellaneum | bellotianum | birrus | bulbiferum | catalaunicum | caulocystidiosum | cavipes | celatum | cinnamomeum | circinans | cistophilum | citrisporum | cohaerens | colvinii | crustuliniforme | cylindrosporum | danicum | discomorbidum | duracinoides | eburneum | echinosporum | erebium | erumpens | excedens | flavidifolium | fuscatum | fusisporum | geminatum | grandisporum | griseopruinatum | harperi | helodes | hiemale | ifeleletorum | immutabile | incarnatulum | indicum | ingratum | ischnostylum | islandicum | kelloggense | lactariolens | laetitiae | laterinum | leucosarx | limbatum | lindae | louiseae | luchuense | luteicystidiatum | lutense | mackinawense | magnicystidiatum | marginatulum | matritense | mediorufum | melleum | mesophaeum | minus | monticola | nanum | nauseosum | naviculosporum | neurophyllum | nigellum | nitidum | nothofagetorum | odoratissimum | olympianum | pallidolabiatum | paludicola | parcivelum | parvicystidiatum | parvisporum | pascuense | perexiguum | plesiocistum | populinum | porphyrosporum | praeolidum | psammophilum | pseudoamarescens | pseudofragilipes | pubescens | pumilum | pungens | pusillum | quercetorum | radicans | radicosoides | radicosum | rostratum | sacchariolens | sagarae | salicicola | sarcophyllum | serratum | sinapizans | sordescens | sordidulum | spetsbergense | sporadicum | subaustrale | subconcolor | subfastibile | subtortum | syrjense | theobrominum | vaccinum | velatum | velutipes | vesterholtii | victoriense | vinosophyllum | viscidissimum | youngii
All related names: aberrans | aeruginosum | affine | agglutinatum | aggregatum | alabamense | alachuanum | albiceps | albidocortinatum | albidocortinatus | albipes | albocolossum | albocrenulatum | albocrenulatus | alvarense | amarellum | angustifolium | angustifolius | angustilamellata | angustispermum | angustisporium | apolectum | apolectus | appendiculatum | aprile | arenicolor | arenosum | argentinum | ascophorus | asperulatum | atrifulvifolium | atrobrunneum | atrosanguineum | augusti | aurantiellum | austroamericanum | bakeri | barrowsii | bellotianus | bicoloratum | bingolense | birra | boulderense | broadwayi | bruchetii | brunneifolium | brunneodiscum | brunneomaculatum | bryogenes | bubalinum | bulbaceum | bulbosum | busporus | caespitosum | californicum | calvinii | calyptrosporum | canariense | candidipes | capniocephalum | capniocephalus | catervarium | catervarius | cerussatum | ceskae | chapmaniae | chlorophyllum | cinchonense | cinereostipes | cinereum | claviceps | clavuligera | clavuligerum | clavulipes | clavus | coarctatum | coarctatus | collariatum | colossus | commune | communis | coniferarum | coprophilum | corrugatum | cortinarioides | crassicystidiatum | crassipes | crassum | crassus | cremeopallidum | cremeum | crustuliniformis | cubense | dasypus | decoratus | deflectens | denigricans | diffracta | diffractum | diffractus | discomorbida | discomorbidus | dissiliens | domardiana | domardianum | dryophilum | dunense | earlei | edule | edulis | edurum | elata | elatellum | elatellus | elatum | elatus | ellipsoideosporium | erebia | erysibodes | evensoniae | exalbidum | exalbidus | exiguifolium | fagicola | fallax | farinaceum | fastibile | fastibilis | favrei | felipponei | felleum | fimbriatus | fimicola | firma | firmum | firmus | flaccidum | flammuloides | flavescens | flavidifolia | flavum | flexuosipes | floridanum | foedatum | fragilior | fragilipes | fragilius | fragrans | fragrantissimum | frenchii | funariophilum | fuscodiscum | fuscodiscus | fuscostipes | fusiformiradicatum | fusiformiradicatus | fusipes | gigaspermum | gigaspora | gigasporum | gigasporus | glabrescens | glutinosa | glutinosum | glutinosus | gomezii | gracile | gregariiforme | gregarium | griseocanescens | griseocanum | griseoscabrosum | griseoscabrosus | griseovelatum | griseum | griseus | groegeri | helvolescens | hemisphaericum | herrmanniae | hesleri | hetieri | holophaea | hormanii | hortense | horticola | hudsoniatum | huijsmaniana | humile | humosum | hydrocybeoides | hygrophilum | idahoense | igneum | ignobile | ignobilis | illicitum | illicitus | imbecillis | incognitum | indecisum | indica | infidum | infidus | insigne | ischnostylus | juneauense | kalmicola | kammala | kanouseae | kauffmanii | kemptoniae | khogianum | kirtonii | kuehneri | lacteocoffeatum | laevatum | laevatus | lamelliconfertum | latericolor | laterinus | lateritium | latifolium | latisporum | laursenii | levisporum | levyanum | lignicolum | limacinum | lindrothii | litoreum | littenii | longicauda | longicaudum | longicaudus | longisporum | lubriciceps | lucidum | lugens | lundqvistii | luteobrunneum | lutescentipes | luteum | macrosporum | maderaspatana | magnimamma | majale | malenconii | mammillatum | mammosum | maritinum | medianum | medianus | mediterranea | mediterraneum | megacarpum | mentiens | mesophaea | mesophaeus | micropyramis | microspora | microsporum | milleri | miserum | mitrata | mitratum | mitratus | montanum | montanus | moseri | mussiva | mussivum | mussivus | mutatum | mutatus | naucorioides | naufragum | naufragus | nauseosus | neerlandica | nigricans | nigromaculatum | nudipes | oblongisporum | obscurum | occidentale | ochraceum | ochroalbidum | octavii | oculatum | odini | odoratissimus | odoratum | olidum | olidus | ollaliense | oregonense | oreophilum | ossa | pallescens | pallida | pallidifolium | pallidoargillaceum | pallidoluctuosum | pallidomarginatum | pallidomarginatus | pallidum | palustre | pamphiliense | parvifructum | parvifructus | patriciae | peckii | perangustisporium | perfarinaceum | perigoense | permollis | perpallidum | perplexum | petiginosum | petiginosus | petrakii | piceicola | pinetorum | pitkinense | platense | polare | politum | porcatosporum | praecaespitosum | praecox | praefarinaceum | praefelleum | praefinitum | praefinitus | praelatifolium | praeviscidum | proletaria | proximum | psammicola | psamminum | psamminus | pseudofastibile | pseudomesophaeum | pseudopunctatum | pseudostrophosum | pudica | pumiloides | punctata | punctatiforme | punctatum | punctatus | pyrophilum | pyrrholepidum | pyrrholepis | queletii | radicatum | radicatus | radicosa | radicosus | remyi | repandum | repandus | riparium | rivulosum | robustum | ruber | rubrofuscum | rubrum | rufosporum | salmonense | sambucinum | sambucinus | sanjuanense | sarcophyllus | scabella | scabellum | senatum | senescens | sericipes | siegelii | siennaecolor | simile | sinapiolens | sinuosa | sinuosum | sinuosus | smithii | sociale | solheimii | sordidulus | sordidum | splendens | spoliatum | spoliatus | squamulosum | stanleyense | stellatosporum | stellatosporus | stenocystis | sterile | sterilis | sterlingii | strophosum | strophosus | suaveolens | subannulatum | subargillaceum | subboreale | subcaespitosum | subcapitatum | subcollariatum | subcollariatus | subfastigiatum | subhepaticum | subincarnatum | sublamellatum | submelinoides | subochracea | subochraceum | subochraceus | subplatense | subrimosum | subrubescens | subsacchariolens | subsaponaceum | subsaponaceus | subscambum | subscambus | substrophosum | subtestaceum | subtestaceus | subumbrinum | subvatricosoides | subvictoriense | subviolaceum | subzonatum | subzonatus | sulcatum | sulcatus | syrjensis | sztoczekii | tenuifolium | termitaria | testacea | testaceum | testaceus | texanum | thomasianum | thomasianus | tomentosum | tomoeae | tortuosum | tortuosus | tottenii | trachysporum | trinidadense | truncatum | truncatus | tumidulum | tumidulus | umbellatum | urbanicola | utahense | vatricosa | vatricosoides | vatricosum | vatricosus | vejlense | velenovskyi | venifer | veniferum | vernale | verrucispora | verruculosum | versipelle | versipellis | victoriae | vinaceogriseum | vinaceoumbrinum | violascens | virgatum | viscidissima | viscidissimus | weberi | wells-kemptoniae | wellsiae | westraliense | xerophilum | zsigmondyi