Published: Nov. Mycol. L. Soucek, Prague: 118 (1939)
Taxonomic status: Heterotypic synonym of H. mesophaeum
Types: CZECH REPUBLIC: Bohemia, SE of Prague, Mencice N of Mnichovice (approx. 49.9104°N, 14.714°E, alt. approx. 360 m a.s.l.) in woodland under Larix sp., Oct. 1934, J. Velenovsky (Holotype. herbarium acc. no. PRM153759, HJB1000154).
Original diagnosis: H. mammilatum, sp. n. Sparsum sed etiam caespitosum, polycephalum, minus, pil. 2-3 cm, semiorbiculari, vertice verruca fusca solida instructo (!), margine diu involuto, glabro, parum hygroph., centro argillaceo, ceterum sordido, juvenili margine albo- fibrilloso. St. 2-3plo longior , 2-3 mm cr., basi fusco non incrass., rigidulus, albus, fibrosus, supra granulosus. Lam. latae, argillaceae, acie albae, late adnatae. Sp. obtuse ovatae 8-10, cyst. magna (25-30) lageniformia, obtusa. Odor debilis. Semper in laricetis pr. Mnichovice autumno ubique frequens. Pilei forma insigne Ex affinitate H. proletarii Vel.
English translation: H. mammillatum sp. n. Scattered but caespitose, with many specimens growing together, small, pileus 2–3 cm, semi-rounded, centre with solid, brown warts, margin long involute, glabrous, weakly hygrophanous, centre clay-coloured, rest sordid, when young with white-fibrillose margin. Stipe 2–3 times longer than diameter, 2–3 mm wide, base brown, not broadened, stiff, white, fibrous, at apex granulose. Lamellae broad, clay-coloured, with white edge, broadly adnate. Spores bluntly ovoid, 8–10 μm. Cystidia large (25–30 μm), lageniform, blunt. Odour weak. In the Larix forests near Mnichovice always everywhere frequent in autumn. The pileus form distinguishes it from H. proletarii.
Commentary: We have examined the holotype. While we have not been able to generate an ITS sequence, our microscopic examination together with the protologue leave us in no doubt that this is: = Hebeloma mesophaeum (Pers.) Quél.
All current species: aanenii | adherens | aestivale | albidulum | alboerumpens | albomarginatum | album | alpinicola | alpinum | ambustiterranum | aminophilum | ammophilum | angelesiense | angustilamellatum | anthracophilum | arcticum | asperosporum | aurantioumbrinum | australe | avellaneum | bellotianum | birrus | bulbiferum | catalaunicum | caulocystidiosum | cavipes | celatum | cinnamomeum | circinans | cistophilum | citrisporum | cohaerens | colvinii | crustuliniforme | cylindrosporum | danicum | discomorbidum | duracinoides | eburneum | echinosporum | erebium | erumpens | excedens | flavidifolium | fuscatum | fusisporum | geminatum | grandisporum | griseopruinatum | harperi | helodes | hiemale | ifeleletorum | immutabile | incarnatulum | indicum | ingratum | ischnostylum | islandicum | kelloggense | lactariolens | laetitiae | laterinum | leucosarx | limbatum | lindae | louiseae | luchuense | luteicystidiatum | lutense | mackinawense | magnicystidiatum | marginatulum | matritense | mediorufum | melleum | mesophaeum | minus | monticola | nanum | nauseosum | naviculosporum | neurophyllum | nigellum | nitidum | nothofagetorum | odoratissimum | olympianum | pallidolabiatum | paludicola | parcivelum | parvicystidiatum | parvisporum | pascuense | perexiguum | plesiocistum | populinum | porphyrosporum | praeolidum | psammophilum | pseudoamarescens | pseudofragilipes | pubescens | pumilum | pungens | pusillum | quercetorum | radicans | radicosoides | radicosum | rostratum | sacchariolens | sagarae | salicicola | sarcophyllum | serratum | sinapizans | sordescens | sordidulum | spetsbergense | sporadicum | subaustrale | subconcolor | subfastibile | subtortum | syrjense | theobrominum | vaccinum | velatum | velutipes | vesterholtii | victoriense | vinosophyllum | viscidissimum | youngii
All related names: aberrans | aeruginosum | affine | agglutinatum | aggregatum | alabamense | alachuanum | albiceps | albidocortinatum | albidocortinatus | albipes | albocolossum | albocrenulatum | albocrenulatus | alvarense | amarellum | angustifolium | angustifolius | angustilamellata | angustispermum | angustisporium | apolectum | apolectus | appendiculatum | aprile | arenicolor | arenosum | argentinum | ascophorus | asperulatum | atrifulvifolium | atrobrunneum | atrosanguineum | augusti | aurantiellum | austroamericanum | bakeri | barrowsii | bellotianus | bicoloratum | bingolense | birra | boulderense | broadwayi | bruchetii | brunneifolium | brunneodiscum | brunneomaculatum | bryogenes | bubalinum | bulbaceum | bulbosum | busporus | caespitosum | californicum | calvinii | calyptrosporum | canariense | candidipes | capniocephalum | capniocephalus | catervarium | catervarius | cerussatum | ceskae | chapmaniae | chlorophyllum | cinchonense | cinereostipes | cinereum | claviceps | clavuligera | clavuligerum | clavulipes | clavus | coarctatum | coarctatus | collariatum | colossus | commune | communis | coniferarum | coprophilum | corrugatum | cortinarioides | crassicystidiatum | crassipes | crassum | crassus | cremeopallidum | cremeum | crustuliniformis | cubense | dasypus | decoratus | deflectens | denigricans | diffracta | diffractum | diffractus | discomorbida | discomorbidus | dissiliens | domardiana | domardianum | dryophilum | dunense | earlei | edule | edulis | edurum | elata | elatellum | elatellus | elatum | elatus | ellipsoideosporium | erebia | erysibodes | evensoniae | exalbidum | exalbidus | exiguifolium | fagicola | fallax | farinaceum | fastibile | fastibilis | favrei | felipponei | felleum | fimbriatus | fimicola | firma | firmum | firmus | flaccidum | flammuloides | flavescens | flavidifolia | flavum | flexuosipes | floridanum | foedatum | fragilior | fragilipes | fragilius | fragrans | fragrantissimum | frenchii | funariophilum | fuscodiscum | fuscodiscus | fuscostipes | fusiformiradicatum | fusiformiradicatus | fusipes | gigaspermum | gigaspora | gigasporum | gigasporus | glabrescens | glutinosa | glutinosum | glutinosus | gomezii | gracile | gregariiforme | gregarium | griseocanescens | griseocanum | griseoscabrosum | griseoscabrosus | griseovelatum | griseum | griseus | groegeri | helvolescens | hemisphaericum | herrmanniae | hesleri | hetieri | holophaea | hormanii | hortense | horticola | hudsoniatum | huijsmaniana | humile | humosum | hydrocybeoides | hygrophilum | idahoense | igneum | ignobile | ignobilis | illicitum | illicitus | imbecillis | incognitum | indecisum | indica | infidum | infidus | insigne | ischnostylus | juneauense | kalmicola | kammala | kanouseae | kauffmanii | kemptoniae | khogianum | kirtonii | kuehneri | lacteocoffeatum | laevatum | laevatus | lamelliconfertum | latericolor | laterinus | lateritium | latifolium | latisporum | laursenii | levisporum | levyanum | lignicolum | limacinum | lindrothii | litoreum | littenii | longicauda | longicaudum | longicaudus | longisporum | lubriciceps | lucidum | lugens | lundqvistii | luteobrunneum | lutescentipes | luteum | macrosporum | maderaspatana | magnimamma | majale | malenconii | mammillatum | mammosum | maritinum | medianum | medianus | mediterranea | mediterraneum | megacarpum | mentiens | mesophaea | mesophaeus | micropyramis | microspora | microsporum | milleri | miserum | mitrata | mitratum | mitratus | montanum | montanus | moseri | mussiva | mussivum | mussivus | mutatum | mutatus | naucorioides | naufragum | naufragus | nauseosus | neerlandica | nigricans | nigromaculatum | nudipes | oblongisporum | obscurum | occidentale | ochraceum | ochroalbidum | octavii | oculatum | odini | odoratissimus | odoratum | olidum | olidus | ollaliense | oregonense | oreophilum | ossa | pallescens | pallida | pallidifolium | pallidoargillaceum | pallidoluctuosum | pallidomarginatum | pallidomarginatus | pallidum | palustre | pamphiliense | parvifructum | parvifructus | patriciae | peckii | perangustisporium | perfarinaceum | perigoense | permollis | perpallidum | perplexum | petiginosum | petiginosus | petrakii | piceicola | pinetorum | pitkinense | platense | polare | politum | porcatosporum | praecaespitosum | praecox | praefarinaceum | praefelleum | praefinitum | praefinitus | praelatifolium | praeviscidum | proletaria | proximum | psammicola | psamminum | psamminus | pseudofastibile | pseudomesophaeum | pseudopunctatum | pseudostrophosum | pudica | pumiloides | punctata | punctatiforme | punctatum | punctatus | pyrophilum | pyrrholepidum | pyrrholepis | queletii | radicatum | radicatus | radicosa | radicosus | remyi | repandum | repandus | riparium | rivulosum | robustum | ruber | rubrofuscum | rubrum | rufosporum | salmonense | sambucinum | sambucinus | sanjuanense | sarcophyllus | scabella | scabellum | senatum | senescens | sericipes | siegelii | siennaecolor | simile | sinapiolens | sinuosa | sinuosum | sinuosus | smithii | sociale | solheimii | sordidulus | sordidum | splendens | spoliatum | spoliatus | squamulosum | stanleyense | stellatosporum | stellatosporus | stenocystis | sterile | sterilis | sterlingii | strophosum | strophosus | suaveolens | subannulatum | subargillaceum | subboreale | subcaespitosum | subcapitatum | subcollariatum | subcollariatus | subfastigiatum | subhepaticum | subincarnatum | sublamellatum | submelinoides | subochracea | subochraceum | subochraceus | subplatense | subrimosum | subrubescens | subsacchariolens | subsaponaceum | subsaponaceus | subscambum | subscambus | substrophosum | subtestaceum | subtestaceus | subumbrinum | subvatricosoides | subvictoriense | subviolaceum | subzonatum | subzonatus | sulcatum | sulcatus | syrjensis | sztoczekii | tenuifolium | termitaria | testacea | testaceum | testaceus | texanum | thomasianum | thomasianus | tomentosum | tomoeae | tortuosum | tortuosus | tottenii | trachysporum | trinidadense | truncatum | truncatus | tumidulum | tumidulus | umbellatum | urbanicola | utahense | vatricosa | vatricosoides | vatricosum | vatricosus | vejlense | velenovskyi | venifer | veniferum | vernale | verrucispora | verruculosum | versipelle | versipellis | victoriae | vinaceogriseum | vinaceoumbrinum | violascens | virgatum | viscidissima | viscidissimus | weberi | wells-kemptoniae | wellsiae | westraliense | xerophilum | zsigmondyi